HomeMy WebLinkAboutAppendix BAPPENDIX B:
STATUS OF DESIGNATED CRITICAL HABITAT: BULL TROUT
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Appendix: B
Status of Designated Critical Habitat: Bull Trout
Past designations of critical habitat have used the terms "primary constituent elements" (PCEs),
"physical and biological features" (PBFs) or "essential features" to characterize the key
components of critical habitat that provide for the conservation of the listed species. The new
critical habitat regulations (81 FR 7214) discontinue use of the terms "PCEs" or "essential
features" and rely exclusively on use of the term PBFs for that purpose because that term is
contained in the statute. To be consistent with that shift in terminology and in recognition that
the terms PBFs, PCEs, and essential habit features are synonymous in meaning, we are only
referring to PBFs herein. Therefore, if a past critical habitat designation defined essential habitat
features or PCEs, they will be referred to as PBFs in this document. This does not change the
approach outlined above for conducting the "destruction or adverse modification" analysis,
which is the same regardless of whether the original designation identified PCEs, PBFs or
essential features.
Current Legal Status of the Critical Habitat
Current Designation
The U.S. Fish and Wildlife Service (Service) published a final critical habitat designation for the
coterminous United States population of the bull trout on October 18, 2010 (USFWS 2010,
entire); the rule became effective on November 17, 2010. A justification document was also
developed to support the rule and is available on the Service's website:
(http://www.f,�vs.gov/pacificibulltrout). The scope of the designation involved the species'
coterminous range, which includes the Coastal, Klamath, Mid -Columbia, Upper Snake,
Columbia Headwaters and St. Mary's Recovery Unit population segments. Rangewide, the
Service designated reservoirs/lakes and stream/shoreline miles as bull trout critical habitat (Table
1). Designated bull trout critical habitat is of two primary use types: 1) spawning and rearing,
and 2) foraging, migration, and overwintering (FMO).
Table 1. Stream/Shoreline Distance and Reservoir/Lake Area Designated as Bull Trout Critical
Habitat.
State
Stream/Shoreline
Miles
Stream/Shoreline
Kilometers
Reservoir/
Lake
Acres
Reservoir/
Lake
Hectares
Idaho
8,771.6
14,116.5
170,217.5
68,884.9
Montana
3,056.5
4,918.9
221,470.7
89,626.4
Nevada
71.8
115.6
-
-
Ore on
2,835.9
4,563.9
30,255.5
12,244.0
Ore on/Idaho
107.7
173.3
-
-
Washington
3,793.3
6,104.8
66,308.1
26,834.0
Washington
(marine)
753.8
1,213.2
-
-
Washington/Idaho
37.2
59.9
-
Washington/Oregon
301.3
484.8
-
-
Total
19,729.0
31,750.8
488,251.7
1 197,589.2
' No shore line is included in Oregon
2 Pine Creek Drainage which falls within Oregon
3 Total of freshwater streams: 18,975
The 2010 revision increases the amount of designated bull trout critical habitat by approximately
76 percent for miles of stream/shoreline and by approximately 71 percent for acres of lakes and
reservoirs compared to the 2005 designation.
The final rule also identifies and designates as critical habitat approximately 1,323.7 km (822.5
miles) of streams/shorelines and 6,758.8 ha (16,701.3 acres) of lakes/reservoirs of unoccupied
habitat to address bull trout conservation needs in specific geographic areas in several areas not
occupied at the time of listing. No unoccupied habitat was included in the 2005 designation.
These unoccupied areas were determined by the Service to be essential for restoring functioning
migratory bull trout populations based on currently available scientific information. These
unoccupied areas often include lower main stem river environments that can provide seasonally
important migration habitat for bull trout. This type of habitat is essential in areas where bull
trout habitat and population loss over time necessitates reestablishing bull trout in currently
unoccupied habitat areas to achieve recovery.
The final rule continues to exclude some critical habitat segments based on a careful balancing of
the benefits of inclusion versus the benefits of exclusion. Critical habitat does not include: 1)
waters adjacent to non -Federal lands covered by legally operative incidental take permits for
habitat conservation plans (HCPs) issued under section 10(a)(1)(B) of the Endangered Species
Act of 1973, as amended (Act), in which bull trout is a covered species on or before the
publication of this final rule; 2) waters within or adjacent to Tribal lands subject to certain
commitments to conserve bull trout or a conservation program that provides aquatic resource
protection and restoration through collaborative efforts, and where the Tribes indicated that
inclusion would impair their relationship with the Service; or 3) waters where impacts to national
security have been identified (USFWS 2010, p. 63903). Excluded areas are approximately 10
percent of the stream/shoreline miles and 4 percent of the lakes and reservoir acreage of
designated critical habitat. Each excluded area is identified in the relevant Critical Habitat Unit
2
(CHU) text, as identified in paragraphs (e)(8) through (e)(41) of the final rule. It is important to
note that the exclusion of waterbodies from designated critical habitat does not negate or
diminish their importance for bull trout conservation. Because exclusions reflect the often
complex pattern of land ownership, designated critical habitat is often fragmented and
interspersed with excluded stream segments.
The Physical and Biological Features
Conservation Role and Description of Critical Habitat
The conservation role of bull trout critical habitat is to support viable core area populations
(USFWS 2010, p. 63898). The core areas reflect the metapopulation structure of bull trout and
are the closest approximation of a biologically functioning unit for the purposes of recovery
planning and risk analyses. CHUB generally encompass one or more core areas and may include
FMO areas, outside of core areas, that are important to the survival and recovery of bull trout.
Thirty-two CHUB within the geographical area occupied by the species at the time of listing are
designated under the revised rule. Twenty-nine of the CHUB contain all of the physical or
biological features identified in this final rule and support multiple life -history requirements.
Three of the mainstem river units in the Columbia and Snake River Basins contain most of the
physical or biological features necessary to support the bull trout's particular use of that habitat,
other than those physical biological features associated with physical and biological features
(PBFs) 5 and 6, which relate to breeding habitat.
The primary function of individual CHUs is to maintain and support core areas, which 1) contain
bull trout populations with the demographic characteristics needed to ensure their persistence and
contain the habitat needed to sustain those characteristics (Rieman and McIntyre 1993, p. 19); 2)
provide for persistence of strong local populations, in part, by providing habitat conditions that
encourage movement of migratory fish (MBTSG 1998, pp. 48-49; Rieman and McIntyre 1993,
pp. 22-23); 3) are large enough to incorporate genetic and phenotypic diversity, but small enough
to ensure connectivity between populations (Hard 1995, pp. 314-315; Healey and Prince 1995, p.
182; MBTSG 1998, pp. 48-49; Rieman and McIntyre 1993, pp. 22-23); and 4) are distributed
throughout the historic range of the species to preserve both genetic and phenotypic adaptations
(Hard 1995, pp. 321-322; MBTSG 1998, pp. 13-16; Rieman and Allendorf 2001, p. 763; Rieman
and McIntyre 1993, p. 23).
Physical and Biological Features for Bull Trout
Within the designated critical habitat areas, the PBFs for bull trout are those habitat components
that are essential for the primary biological needs of foraging, reproducing, rearing of young,
dispersal, genetic exchange, or sheltering. Based on our current knowledge of the life history,
biology, and ecology of this species and the characteristics of the habitat necessary to sustain its
essential life -history functions, we have determined that the PBFs, as described within USFWS
2010, are essential for the conservation of bull trout. A summary of those PBFs follows.
1. Springs, seeps, groundwater sources, and subsurface water connectivity (hyporheic
flows) to contribute to water quality and quantity and provide thermal refugia.
2. Migration habitats with minimal physical, biological, or water quality impediments
between spawning, rearing, overwintering, and freshwater and marine foraging habitats,
including but not limited to permanent, partial, intermittent, or seasonal barriers.
3. An abundant food base, including terrestrial organisms of riparian origin, aquatic
macroinvertebrates, and forage fish.
4. Complex river, stream, lake, reservoir, and marine shoreline aquatic environments, and
processes that establish and maintain these aquatic environments, with features such as
large wood, side channels, pools, undercut banks and unembedded substrates, to provide
a variety of depths, gradients, velocities, and structure.
5. Water temperatures ranging from 2 °C to 15 °C, with adequate thermal refugia available
for temperatures that exceed the upper end of this range. Specific temperatures within
this range will depend on bull trout life -history stage and form; geography; elevation;
diurnal and seasonal variation; shading, such as that provided by riparian habitat;
streamflow; and local groundwater influence.
6. In spawning and rearing areas, substrate of sufficient amount, size, and composition to
ensure success of egg and embryo overwinter survival, fry emergence, and young -of -the -
year and juvenile survival. A minimal amount of fine sediment, generally ranging in size
from silt to coarse sand, embedded in larger substrates, is characteristic of these
conditions. The size and amounts of fine sediment suitable to bull trout will likely vary
from system to system.
7. A natural hydrograph, including peak, high, low, and base flows within historic and
seasonal ranges or, if flows are controlled, minimal flow departure from a natural
hydrograph.
8. Sufficient water quality and quantity such that normal reproduction, growth, and survival
are not inhibited.
9. Sufficiently low levels of occurrence of non-native predatory (e.g., lake trout, walleye,
northern pike, smallmouth bass); interbreeding (e.g., brook trout); or competing (e.g.,
brown trout) species that, if present, are adequately temporally and spatially isolated from
bull trout.
The revised PBF's are similar to those previously in effect under the 2005 designation. The most
significant modification is the addition of a ninth PBF to address the presence of nonnative
predatory or competitive fish species. Although this PBF applies to both the freshwater and
marine environments, currently no non-native fish species are of concern in the marine
environment, though this could change in the future.
Note that only PBFs 2, 3, 4, 5, and 8 apply to marine nearshore waters identified as critical
habitat. Also, lakes and reservoirs within the CHUs also contain most of the physical or
biological features necessary to support bull trout, with the exception of those associated with
PBFs 1 and 6. Additionally, all except PBF 6 apply to FMO habitat designated as critical
habitat.
2
Critical habitat includes the stream channels within the designated stream reaches and has a
lateral extent as defined by the bankfull elevation on one bank to the bankfull elevation on the
opposite bank. Bankfull elevation is the level at which water begins to leave the channel and
move into the floodplain and is reached at a discharge that generally has a recurrence interval of
1 to 2 years on the annual flood series. If bankfull elevation is not evident on either bank, the
ordinary high-water line must be used to determine the lateral extent of critical habitat. The
lateral extent of designated lakes is defined by the perimeter of the waterbody as mapped on
standard 1:24,000 scale topographic maps. The Service assumes in many cases this is the full -
pool level of the waterbody. In areas where only one side of the waterbody is designated (where
only one side is excluded), the mid -line of the waterbody represents the lateral extent of critical
habitat.
In marine nearshore areas, the inshore extent of critical habitat is the mean higher high-water
(MHHW) line, including the uppermost reach of the saltwater wedge within tidally influenced
freshwater heads of estuaries. The MHHW line refers to the average of all the higher high-water
heights of the two daily tidal levels. Marine critical habitat extends offshore to the depth of 10
meters (m) (33 ft) relative to the mean low low-water (MLLW) line (zero tidal level or average
of all the lower low-water heights of the two daily tidal levels). This area between the MHHW
line and minus 10 in MLLW line (the average extent of the photic zone) is considered the habitat
most consistently used by bull trout in marine waters based on known use, forage fish
availability, and ongoing migration studies and captures geological and ecological processes
important to maintaining these habitats. This area contains essential foraging habitat and
migration corridors such as estuaries, bays, inlets, shallow subtidal areas, and intertidal flats.
Adjacent shoreline riparian areas, bluffs, and uplands are not designated as critical habitat.
However, it should be recognized that the quality of marine and freshwater habitat along streams,
lakes, and shorelines is intrinsically related to the character of these adjacent features, and that
human activities that occur outside of the designated critical habitat can have major effects on
physical and biological features of the aquatic environment.
Activities that cause adverse effects to critical habitat are evaluated to determine if they are
likely to "destroy or adversely modify" critical habitat by no longer serving the intended
conservation role for the species or retaining those PBFs that relate to the ability of the area to at
least periodically support the species. Activities that may destroy or adversely modify critical
habitat are those that alter the PBFs to such an extent that the conservation value of critical
habitat is appreciably reduced (USFWS 2010, pp. 63898:63943; USFWS 2004a, pp. 140-193;
USFWS 2004b, pp. 69-114). The Service's evaluation must be conducted at the scale of the
entire critical habitat area designated, unless otherwise stated in the final critical habitat rule
(USFWS and NMFS 1998, Ch. 4 p. 39). Thus, adverse modification of bull trout critical habitat
is evaluated at the scale of the final designation, which includes the critical habitat designated for
the Klamath River, Jarbidge River, Columbia River, Coastal -Puget Sound, and Saint Mary -Belly
River population segments. However, we consider all 32 CHUs to contain features or areas
essential to the conservation of the bull trout (USFWS 2010, pp. 63898:63901, 63944).
Therefore, if a proposed action would alter the physical or biological features of critical habitat
to an extent that appreciably reduces the conservation function of one or more critical habitat
units for bull trout, a finding of adverse modification of the entire designated critical habitat area
may be warranted (USFWS 2010, pp. 63898:63943).
Current Critical Habitat Condition Rangewide
The condition of bull trout critical habitat varies across its range from poor to good. Although
still relatively widely distributed across its historic range, the bull trout occurs in low numbers in
many areas, and populations are considered depressed or declining across much of its range
(Ratliff and Howell 1992, entire; Schill 1992, p. 40; Thomas 1992, p. 28; Buchanan et al. 1997,
p. vii; Rieman et al. 1997, pp. 15-16; Quigley and Arbelbide 1997, pp. 1176-1177). This
condition reflects the condition of bull trout habitat. The decline of bull trout is primarily due to
habitat degradation and fragmentation, blockage of migratory corridors, poor water quality, past
fisheries management practices, impoundments, dams, water diversions, and the introduction of
nonnative species (USFWS 1998, pp. 31648-31649; USFWS 1999, p. 17111).
There is widespread agreement in the scientific literature that many factors related to human
activities have impacted bull trout and their habitat, and continue to do so. Among the many
factors that contribute to degraded PBFs, those which appear to be particularly significant and
have resulted in a legacy of degraded habitat conditions are as follows: 1) fragmentation and
isolation of local populations due to the proliferation of dams and water diversions that have
eliminated habitat, altered water flow and temperature regimes, and impeded migratory
movements (Dunham and Rieman 1999, p. 652; Rieman and McIntyre 1993, p. 7); 2)
degradation of spawning and rearing habitat and upper watershed areas, particularly alterations
in sedimentation rates and water temperature, resulting from forest and rangeland practices and
intensive development of roads (Fraley and Shepard 1989, p. 141; MBTSG 1998, pp. ii - v, 20-
45); 3) the introduction and spread of nonnative fish species, particularly brook trout and lake
trout, as a result of fish stocking and degraded habitat conditions, which compete with bull trout
for limited resources and, in the case of brook trout, hybridize with bull trout (Leary et al. 1993,
p. 857; Rieman et al. 2006, pp. 73-76); 4) in the Coastal -Puget Sound region where
amphidromous bull trout occur, degradation of mainstem river FMO habitat, and the degradation
and loss of marine nearshore foraging and migration habitat due to urban and residential
development; and 5) degradation of FMO habitat resulting from reduced prey base, roads,
agriculture, development, and dams.
Effects of Climate Change on Bull Trout Critical Habitat
One objective of the final rule was to identify and protect those habitats that provide resiliency
for bull trout use in the face of climate change. Over a period of decades, climate change may
directly threaten the integrity of the essential physical or biological features described in PBFs 1,
2, 3, 5, 7, 8, and 9. Protecting bull trout strongholds and cold water refugia from disturbance
and ensuring connectivity among populations were important considerations in addressing this
potential impact. Additionally, climate change may exacerbate habitat degradation impacts both
physically (e.g., decreased base flows, increased water temperatures) and biologically (e.g.,
increased competition with non-native fishes).
Many of the PBFs for bull trout may be affected by the presence of toxics and/or increased water
temperatures within the environment. The effects will vary greatly depending on a number of
factors which include which toxic substance is present, the amount of temperature increase, the
likelihood that critical habitat would be affected (probability), and the severity and intensity of
any effects that might occur (magnitude).
0
The ability to assign the effects of gradual global climate change bull trout critical habitat or to a
specific location on the ground is beyond our technical capabilities at this time.
LITERATURE CITED
Buchanan, D.V., M.L. Hanson, and R.M. Hooton. 1997. Status of Oregon's bull trout. Oregon
Department of Fish and Wildlife. 168 pp.
Dunham, J.B. and B.E. Rieman. 1999. Metapopulation structure of bull trout: Influences of
physical, biotic, and geometrical landscape characteristics. Ecological Applications
9:642-655. 15 pp.
Fraley, J.J., and B.B. Shepard. 1989. Life history, ecology and population status of migratory
bull trout (Salvelinus confluentus) in the Flathead Lake and River System, Montana.
Northwest Science 63(4):133-143.
Hard, J. 1995. A quantitative genetic perspective on the conservation of intraspecific diversity.
American Fisheries Society Symposium 17: 304-326. 22 pp.
Healey, M.C. and A. Prince. 1995. Scales of variation in life history tactics of Pacific salmon
and the conservation of phenotype and genotype. American Fisheries Society
Symposium 17:176-84. 10 pp.
Leary, R.F., F.W. Allendorf, and S.H. Forbes. 1993. Conservation genetics of bull trout in the
Columbia and Klamath River drainages. Conservation Biology [CONSERV. BIOL.]
7:856-865.
MBTSG (Montana Bull Trout Scientific Group). 1998. The relationship between land
management activities and habitat requirements of bull trout. Prepared for Montana Bull
Trout Restoration Team. Helena, Montana. 86 pp.
Quigley, T.M., and S.J. Arbelbide, tech. eds. 1997. An assessment of ecosystem components in
the Interior Columbia Basin and portions of the Klamath and Great Basins: volume I11.
Gen. Tech. Rep. PNW-GTR-405. Portland, OR: U.S. Department of Agriculture, Forest
Service, Pacific Northwest Research Station. 4 vol. 13 pp.
Ratliff, D.E., and P.J. Howell. 1992. The status of bull trout populations in Oregon. Pages 10-
17 in: P.J. Howell and D.V. Buchanan (eds). Proceedings of the Gearhart Mountain bull
trout workshop. Oregon Chapter of the American Fisheries Society, Corvallis. 8 pp.
Rieman, B.E., and J.D. McIntyre. 1993. Demographic and habitat requirements of bull trout
Salvelinus confluentus. General Technical Report INT -GTR- 302. U.S. Department of
Agriculture, Forest Service, Intermountain Research Station, Ogden, Utah. 42 pp.
Rieman, B.E., and F.W. Allendorf. 2001. Effective population size and genetic conservation
criteria for bull trout. North American Journal of Fisheries Management 21:756-764.
American Fisheries Society, Bethesda, Maryland. 10 pp.
7
Rieman, B.E., D.C. Lee and R.F. Thurow. 1997. Distribution, status and likely future trends of
Bull trout within the Columbia River and Klamath River basins. North American Journal
of Fisheries Management 17:1111-1125.48 pp.
Rieman, B.E., J.T. Peterson and D.L. Myers. 2006. Have brook trout (Salvelinus fontinalis)
displaced bull trout (Salvelinus confluentus) along longitudinal gradients in central Idaho
streams? Canadian Journal of Fisheries and Aquatic Sciences. Vol. 63, No. 1, pp. 63-78.
16 pp.
Schill, D.J. 1992. River and stream investigations. Job Performance Report, Project F -73-R-13.
Idaho Department of Fish and Game, Boise, Idaho. 66 pp.
Thomas, G. 1992. Status of bull trout in Montana. Report prepared for Montana Department of
Fish, Wildlife and Parks, Helena, Montana. 108 pp.
USFWS (U.S. Fish and Wildlife Service) and NMFS (National Marine Fisheries Service). 1998.
Consultation handbook: procedures for conducting consultation and conference activities
under Section 7 of the Endangered Species Act. 315pp.
USFWS (U.S. Fish and Wildlife Service). 1998. Determination of threatened status for the
Klamath River and Columbia River distinct population segments of bull trout. Federal
Register Vol. 63 31647-31674. 28 pp.
. 1999. Determination of threatened status for bull trout for the Jarbidge River population
segment of bull trout. Federal Register Vol. 64 17110-17125. 16 pp.
. 2004a. Draft Recovery Plan for the Coastal -Puget Sound Distinct Population Segment
of Bull Trout (Salvelinus confluentus). U.S. Fish and Wildlife Service, Portland, Oregon.
297 pp.
. 2004b. Draft Recovery Plan for the Jarbidge Distinct Population Segment of Bull Trout
(Salvelinus confluentus). U.S. Fish and Wildlife Service, Portland, Oregon. 148 pp.
. 2010. Revised designation of critical habitat for bull trout in the coterminous United
States. Federal Register Vol 75, No. 200. 63898-64070.